For over 750 years these have been kept here at the church of St. Francis (XIII cent.), at Cortona (near Florence).
This is the second largest church in the world dedicated to St. Francis, after his Basilica in Assisi (both were designed and constructed by Friar Elias, follower and companion of St. Francis, his Vicar and subsequently Minister General of the Order). With its gothic nave and trussed roof, this style became the model for Franciscan churches. Friar Elias himself carried these relics here from Assisi.